Epirus: This little-known region of the Greek mainland has the world’s deepest gorge, a landscape like Scotland’s and mountain villages to rival those of Provence, says Caroline Shearing. Photograph by Andreas Athan
Tadpoles overhead. Overall winner, Category winner: Ecology and Environmental Science. Tadpoles of many anuran species come in high numbers, but not many make it to adulthood. Here a group of common toad (Bufo bufo) tadpoles is seen from below by Bert Willaert, Belgium. It was shot with a canon G12 camera in a Recsea underwater housing and a Dyron dome port.